Ecology, Evolution, and Organismal Biology (EEOB)

EEOB 507. Advanced Animal Behavior.

(3-0) Cr. 3. S. Prereq: Graduate standing, BIOL 354, or permission of instructor
Analysis of current research in animal behavior. Topics covered may include behavioral ecology, mechanisms of behavior, evolution of behavior, applications of animal behavior to conservation biology, and applications of animal behavior to wild animals in captivity.

EEOB 514. Life History and Reproductive Strategies.

(Dual-listed with BIOL 414). (3-0) Cr. 3. F. Prereq: BIOL 315 or equivalent; course in population ecology suggested.
Evolution of ecological adaptations at the individual, population, community and landscape levels. Emphasis is on evolutionary mechanisms and adaptive strategies; units and mechanisms of evolution, life history strategies, species interactions and organization of communities, behavior, and patterns of distribution, speciation and macroevolution.

EEOB 531. Conservation Biology.

(Cross-listed with A ECL). (3-0) Cr. 3. Alt. S., offered even-numbered years. Prereq: BIOL 312; BIOL 313 or graduate standing
Examination of conservation issues from a population and a community perspective. Population-level analysis will focus on the role of genetics, demography, and environment in determining population viability. Community perspectives will focus on topics such as habitat fragmentation, reserve design, biodiversity assessment, and restoration ecology.

EEOB 531I. Conservation Biology.

(Cross-listed with A ECL, IA LL). Cr. 4. Alt. SS., offered even-numbered years. Prereq: IA LL 312I
Population-and community-level examination of factors influencing the viability of plant and animal populations from both demographic and genetic perspectives; assessment of biodiversity; design and management of preserves.

EEOB 534. Endocrinology.

(Dual-listed with BIOL 434). (3-0) Cr. 3. S. Prereq: BIOL 211, BIOL 212
Chemical integration of vertebrate organisms. The structure, development, and evolution of the endocrine glands and the function and structure of their hormones.

EEOB 535. Restoration Ecology.

(Cross-listed with ENSCI, NREM). (2-3) Cr. 3. F. Prereq: BIOL 366 or BIOL 474 or graduate standing
Theory and practice of restoring animal and plant diversity, structure and function of disturbed ecosystems. Restored freshwater wetlands, forests, prairies and reintroduced species populations will be used as case studies.

EEOB 535I. Restoration Ecology.

(Cross-listed with A ECL, ENSCI, IA LL). Cr. 4. Alt. SS., offered even-numbered years. Prereq: A course in ecology
Ecological principles for the restoration of native ecosystems; establishment (site preparation, selection of seed mixes, planting techniques) and management (fire, mowing, weed control) of native vegetation; evaluation of restorations. Emphasis on the restoration of prairie and wetland vegetation.

EEOB 539. Environmental Physiology.

(Dual-listed with BIOL 439). (3-3) Cr. 3-4. Alt. S., offered even-numbered years. Prereq: BIOL 335 or A ECL 311, physics recommended
Graduate study in conjunction with BIOL 439. Physiological adaptations to the environment with emphasis on vertebrates.

EEOB 542. Introduction to Molecular Biology Techniques.

(Cross-listed with B M S, FS HN, GDCB, HORT, NREM, NUTRS, V MPM, VDPAM). Cr. 1. Repeatable. F.S.SS. Prereq: Graduate classification
Sessions in basic molecular biology techniques and related procedures. Offered on a satisfactory-fail basis only.

EEOB 542A. Introduction to Molecular Biology Techniques: DNA Techniques.

(Cross-listed with B M S, BBMB, FS HN, GDCB, HORT, NREM, NUTRS, V MPM, VDPAM). Cr. 1. Repeatable. F.S.SS. Prereq: Graduate classification
Includes genetic engineering procedures, sequencing, PCR, and genotyping. Offered on a satisfactory-fail basis only.

EEOB 542C. Introduction to Molecular Biology Techniques: Cell Techniques.

(Cross-listed with B M S, BBMB, FS HN, GDCB, HORT, NREM, NUTRS, V MPM, VDPAM). Cr. 1. Repeatable. F.S. Prereq: Graduate classification
Includes immunophenotyping, ELISA, flow cytometry, microscopic techniques, image analysis, confocal, multiphoton and laser capture microdissection. Offered on a satisfactory-fail basis only.

EEOB 542D. Introduction to Molecular Biology Techniques: Plant Transformation.

(Cross-listed with B M S, FS HN, GDCB, HORT, NREM, NUTRS, V MPM, VDPAM). Cr. 1. Repeatable. S. Prereq: Graduate classification
Includes Agrobacterium and particle gun-mediated transformation of tobacco, Arabidopsis, and maize, and analysis of tranformants. Offered on a satisfactory-fail basis only.

EEOB 542E. Introduction to Molecular Biology Techniques: Proteomics.

(Cross-listed with B M S, BBMB, FS HN, GDCB, HORT, NREM, NUTRS, V MPM, VDPAM). Cr. 1. Repeatable. F. Prereq: Graduate classification
Includes two-dimensional electrophoresis, laser scanning, mass spectrometry, and database searching. Offered on a satisfactory-fail basis only.

EEOB 542F. Introduction to Molecular Biology Techniques: Metabolomics.

(Cross-listed with B M S, BBMB, FS HN, GDCB, HORT, NREM, NUTRS, V MPM, VDPAM). Cr. 1. Repeatable. F. Prereq: Graduate classification
Includes metabolomics and the techniques involved in metabolite profiling. For non-chemistry majoring students who are seeking analytical aspects into their biological research projects. Offered on a satisfactory-fail basis only.

EEOB 542G. Introduction to Molecular Biology Techniques: Genomic.

(Cross-listed with B M S, FS HN, GDCB, HORT, NREM, NUTRS, V MPM, VDPAM). Cr. 1. Repeatable. S. Prereq: Graduate classification
Offered on a satisfactory-fail basis only.

EEOB 544. Introduction to Bioinformatics.

(Dual-listed with BIOL 444). (4-0) Cr. 4. F. Prereq: MATH 165 or STAT 401 or equivalent
Broad overview of bioinformatics with a significant problem-solving component, including hands-on practice using computational tools to solve a variety of biological problems. Topics include: database searching, sequence alignment, gene prediction, RNA and protein structure prediction, construction of phylogenetic trees, comparative and functional genomics, systems biology.

EEOB 551. Plant Evolution and Phylogeny.

(Dual-listed with BIOL 451). (3-3) Cr. 4. F. Prereq: BIOL 315 or equivalent.
Survey of land plant evolution; phylogenetic comparison of anatomical, reproductive, and life history specializations. Relationships among bryophytes, lycophytes, pteridophytes, gymnosperms, and angiosperms emphasizing significant evolutionary changes documented by paleobotanical, morphological, and molecular studies.

EEOB 553. Agrostology.

(2-3) Cr. 3. Alt. F., offered even-numbered years. Prereq: BIOL 366
Structure, identification, classification, phylogeny, and economic aspects of grasses and related families.

EEOB 555. Bryophyte and Lichen Biodiversity.

(Dual-listed with BIOL 455). Cr. 3. Prereq: BIOL 212, BIOL 212L
Introduction to the biology and ecology of mosses, liverworts, and lichens. Emphasis on identification and diversity of local representatives of these three groups of organisms. Required field trips and service-learning.

EEOB 559. Mammalogy.

(2-3) Cr. 3. S. Prereq: BIOL 351 or A ECL 365
Biology, ecology, and evolution of mammals. Emphasis on structure, physiological adaptation to different environments, behavior, reproduction, roles of mammals in ecosystems, and conservation. Laboratory focus on identification, distribution, habits, and habitats of mammals.

EEOB 560. Resource Ecology.

(2-3) Cr. 3. Alt. S., offered even-numbered years. Prereq: BIOL 212, BIOL 212L, BIOL 312; STAT 101 or STAT 104 or graduate standing
Ecological and economical management of sustainable biological resources. Unifying current management concepts and models in wildlife, fisheries, water quality, forestry, recreation, and agriculture. Research problems.

EEOB 562. Evolutionary Genetics.

(3-0) Cr. 3. Alt. S., offered odd-numbered years. Prereq: Permission of instructor
Seminar/discussion course covering the genetic basis of evolutionary processes in multicellular organisms.

EEOB 563. Molecular Phylogenetics.

(2-3) Cr. 3. F. Prereq: BIOL 313 and BIOL 315
An overview of the theory underlying phylogenetic analysis and the application of phylogenetic methods to molecular datasets. The course emphasizes a hands-on approach to molecular phylogenetics and combines lecture presentations with computer exercises and discussion of original scientific literature.

EEOB 564. Wetland Ecology.

(Cross-listed with ENSCI). (3-0) Cr. 3. S. Prereq: 15 credits in biological sciences
Ecology, classification, creation and restoration, and management of wetlands. Emphasis on North American temperate wetlands.

EEOB 564I. Wetland Ecology.

(Cross-listed with ENSCI, IA LL). Cr. 4. SS. Prereq: IA LL 312I
Ecology, classification, creation, restoration, and management of wetlands. Field studies will examine the composition, structure and functions of local natural wetlands and restored prairie pothole wetlands. Individual or group projects.

EEOB 565. Morphometric Analysis.

(Dual-listed with BIOL 465). (3-2) Cr. 4. Alt. S., offered even-numbered years. Prereq: STAT 401
A comprehensive overview of the theory and methods for the analysis of biological shape with emphasis on data acquisition, standardization, statistical analysis, and visualization of results. Methods for both landmark and outline data will be discussed.

EEOB 566. Molecular Evolution.

(3-0) Cr. 3. Alt. F., offered even-numbered years. Prereq: Permission of instructor
Seminar/discussion course covering the fundamentals of molecular evolution. Emphasis is placed on original scientific literature and current topics, including rates and patterns of genetic divergence; nucleotide and allelic diversity; molecular clocks; gene duplications; genome structure; organellar genomes; polyploidy; transposable elements; and modes and mechanisms of gene and genome evolution.

EEOB 567. Empirical Population Genetics.

(3-0) Cr. 3. F. Prereq: Permission of instructor
An overview of fundamental population genetic theory and the ecological and evolutionary factors underlying the distribution of genetic variation within and among natural populations. Emphasis on the analysis of inbreeding, breeding systems, parentage, relatedness, spatial autocorrelation, effective population size, hierarchial population models, and phylogeography.

EEOB 568. Advanced Systematics.

(Cross-listed with ENT). (2-3) Cr. 3. Alt. S., offered odd-numbered years. Prereq: Permission of instructor
Principles and practice of systematic biology; taxonomy, nomenclature and classification of plants and animals; sources and interpretation of systematic data; speciation; fundamentals of phylogenetic systematics.

EEOB 569. Biogeography.

(3-0) Cr. 3. Alt. S., offered even-numbered years. Prereq: BIOL 315 or equivalent; permission of instructor
Principles underlying the geographic distribution of organisms throughout the world; biological influences of geological history and tectonic movements; role of climate, migration, dispersal, habitat, and phylogeny on past and present organismal distribution patterns; biogeographic methods.

EEOB 570. Landscape Ecology.

(Cross-listed with A ECL). (2-3) Cr. 3. Alt. F., offered even-numbered years. Prereq: Permission of instructor; EEOB 588; a course in calculus
The study of ecological and evolutionary processes within a spatial context with emphasis on behavior, population, and community dynamics.

EEOB 573. Techniques for Biology Teaching.

(Cross-listed with A ECL, IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573A. Techniques for Biology Teaching : Animal Biology.

(Cross-listed with A ECL, IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573B. Techniques for Biology Teaching: Plant Biology.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573C. Techniques for Biology Teaching: Fungi and Lichens.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573D. Techniques for Biology Teaching: Aquatic Ecology.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573E. Techniques for Biology Teaching: Prairie Ecology.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573F. Techniques for Biology Teaching: Wetland Ecology.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573G. Techniques for Biology Teaching: Limnology.

(Cross-listed with A ECL, IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573H. Techniques for Biology Teaching: Animal Behavior.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573I. Techniques for Biology Teaching: Insect Ecology.

(Cross-listed with A ECL, IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573J. Techniques for Biology Teaching: Biology of Invertebrates.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573K. Techniques for Biology Teaching: Non-invasive Use of Living Organisms.

(Cross-listed with IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 573W. Techniques for Biology Teaching: Project WET.

(Cross-listed with A ECL, IA LL). Cr. 1-2. Repeatable. SS.
The development and implementation of laboratory exercises suitable for inclusion in elementary, middle, high school, and community college biology and environmental courses. Exercises will be built around common organisms and ecosystems in Iowa. Field trips.

EEOB 575I. Field Mycology.

(Cross-listed with IA LL). Cr. 4. Alt. SS., offered even-numbered years.
Identification and classification of the common fungi; techniques for identification, preservation, and culture practiced with members of the various fungi groups.

EEOB 576. Functional Ecology.

(Dual-listed with BIOL 476). (3-0) Cr. 3. Alt. S., offered odd-numbered years. Prereq: BIOL 312
The nature of adaptations to physical and biotic environments. Biophysical, biomechanical, and physiological bases of the structure, form, growth, distribution, and abundance of organisms.

EEOB 577. Concepts in Theoretical Ecology and Evolution.

(2-0) Cr. 1. Alt. F., offered even-numbered years.
Readings and discussion of influential ideas in ecological and evolutionary theory, with an emphasis on how models are used as conceptual tools for building synthetic paradigms. Topics are chosen according to student interests; may include spatial ecology, behavioral theory, chaos, community assembly and biodiversity, and others.

EEOB 578. Foundations of Theoretical Ecology and Evolution.

(3-0) Cr. 3. Alt. S., offered even-numbered years. Prereq: 1 semester of calculus or permission of instructor.
Quantitative exploration of classic models and results in ecological and evolutionary theory. Introduction to conceptual, mathematical, and programming tools needed to build and analyze models.

EEOB 580I. Ecology and Systematics of Diatoms.

(Cross-listed with IA LL). Cr. 4. SS.
Field and laboratory study of freshwater diatoms; techniques in collection, preparation, and identification of diatom samples; study of environmental factors affecting growth, distribution, taxonomic characters; project design and execution including construction of reference and voucher collections and data organization and analysis.

EEOB 581. Environmental Systems I: Introduction to Environmental Systems.

(Dual-listed with BIOL 381). (Cross-listed with ENSCI). Cr. 3-4. F. Prereq: 12 credits of natural science including biology and chemistry
Introduction to the structure and function of natural environmental systems. Emphasis on the analysis of material and energy flows in natural environmental systems and the primary environmental factors controlling these systems.

EEOB 582. Environmental Systems II: Analysis of Environmental Systems.

(Dual-listed with BIOL 382). (Cross-listed with ENSCI). (2-2) Cr. 3. S. Prereq: ENSCI 581
Continuation of ENSCI 581. Systems approach to the analysis of material and energy flows in natural environmental systems and the primary environmental factors controlling these systems.

EEOB 584. Advanced Ecosystem Ecology.

(Cross-listed with ENSCI). (3-0) Cr. 3. Alt. S., offered even-numbered years. Prereq: Combined 12 credits in biology, chemistry, and physics.
Advanced studies of ecosystems and the biological and physical factors that influence their properties and dynamics. Conceptual foundations and modern approaches to ecosystem studies. Interactions among organisms, biological diversity, and ecosystem attributes. Quantitative analyses of accumulations, transformations, and fluxes of nutrients, water, and energy within and among ecosystems. Global change issues.

EEOB 585. Advanced Community Ecology.

(2-3) Cr. 3. Alt. F., offered even-numbered years. Prereq: BIOL 312
Factors controlling species diversity, species abundance, and the structure and function of communities in space and time. Relationships between species diversity and ecosystem process rates and community stability.

EEOB 586. Aquatic Ecology.

(Dual-listed with BIOL 486). (Cross-listed with ENSCI). (3-0) Cr. 3. F. Prereq: BIOL 312 or ENSCI 381 or ENSCI 402 or NREM 301
Structure and function of aquatic ecosystems with application to fishery and pollution problems. Emphasis on lacustrine, riverine, and wetland ecology.

EEOB 586L. Aquatic Ecology Laboratory.

(Dual-listed with BIOL 486L). (Cross-listed with ENSCI). (0-3) Cr. 1. F. Prereq: Concurrent enrollment in BIOL 486
Field trips and laboratory exercises to accompany 486. Hands-on experience with aquatic research and monitoring techniques and concepts.

EEOB 587. Microbial Ecology.

(Dual-listed with BIOL 487). (Cross-listed with ENSCI, MICRO). (3-0) Cr. 3. F. Prereq: Six credits in biology and 6 credits in chemistry
Introduction to major functional groups of autotrophic and heterotrophic microorganisms and their roles in natural systems.

EEOB 589. Population Ecology.

(Dual-listed with BIOL 489). (Cross-listed with A ECL). (2-2) Cr. 3. F. Prereq: BIOL 312, STAT 101 or STAT 104, a course in calculus, or graduate standing
Concepts and theories of population dynamics with emphasis on models of growth, predation, competition, and regulation.

EEOB 590. Graduate Independent Study.

(Cross-listed with A ECL, ANTHR, IA LL). Cr. 1-4. Repeatable. SS. Prereq: Graduate classification and permission of instructor

EEOB 590A. Special Topics: Current Topics in Ecology.

Cr. 1-3. Repeatable. Prereq: 10 credits in biology, permission of instructor

EEOB 590B. Special Topics: Current Topics in Evolutionary Biology.

Cr. 1-3. Repeatable. Prereq: 10 credits in biology, permission of instructor

EEOB 590C. Special Topics: Current Topics in Organismal Biology.

Cr. 1-3. Repeatable. Prereq: 10 credits in biology, permission of instructor

EEOB 590I. Special Topics: Graduate Independent Study.

(Cross-listed with A ECL, ANTHR, IA LL). Cr. 1-4. Repeatable. SS. Prereq: Graduate classification and permission of instructor

EEOB 596. Ecology and Society.

(3-0) Cr. 3. Prereq: Graduate classification in biological or environmental sciences/studies with at least one course in ecology
Analysis of conceptual and methodological debates in ecology. Historical development of competing research traditions and philosophies. Topics include i) methodological issues in ecological science, ii) conceptual issues in theoretical ecology, iii) conceptual issues in applied ecology, iv) relation of ecology to environmental and social issues.

EEOB 599. Creative Component.

Cr. arr.
Research toward nonthesis master's degree.

EEOB 611. Analysis of Populations.

(Cross-listed with A ECL). (2-2) Cr. 3. Alt. F., offered odd-numbered years. Prereq: BIOL 312; STAT 401; a course in calculus
Quantitative techniques for analyzing vertebrate population data to estimate parameters such as density and survival. Emphasis on statistical inference and computing.

EEOB 679. Light Microscopy.

(Cross-listed with GDCB, MICRO). (2-9) Cr. 5. Prereq: Permission of instructor
Current theories encompassing light optics and their applications for specimen preservation, paraffin and resin sectioning, general staining, histochemistry, cytophotometry, immunocytochemistry, autoradiography, image digitization, processing and presentation, and digital macro- and micrography. Limit of 10 students.

EEOB 680. Scanning Electron Microscopy.

(Cross-listed with GDCB, MICRO). (2-9) Cr. 5. Prereq: Permission of instructor
Current theories encompassing scanning electron optics and their applications for high and low vacuum microscopy, specimen chemical and cryopreservation methods, x-ray microanalysis, backscattered and topographic imaging, image digitization, processing and presentation. Limit of 10 students.

EEOB 681. Transmission Electron Microscopy.

(Cross-listed with GDCB, MICRO). (2-9) Cr. 5. Prereq: GDCB 679 and permission of instructor
Current theories encompassing electron optics and their applications for chemical and physical specimen preservation, ultramicrotomy, general staining and cytochemistry, immunocytochemistry, autoradiography, negative staining and shadowing, x-ray microanalysis, image digitization, processing and presentation.

EEOB 698. Seminar.

Cr. 1. Repeatable.
Meetings of graduate students and faculty to discuss recent literature and problems under investigation.

EEOB 699. Research.

Cr. arr. Repeatable.
Research for thesis or dissertation. Offered on a satisfactory-fail basis only.

EEOB 699I. Research.

(Cross-listed with A ECL, ANTHR, GDCB, IA LL). Cr. 1-4. Repeatable.